David Reynolds

Postdoctoral Research Associate I

David’s research  focuses on the development and application of novel oceanographic records derived from variations in the width and geochemical (d18O, d13C and 14C) composition of annual growth increments formed in the shells of long-lived marine bivalve molluscs. These records, known as sclerochronologies, are regarded as the marine and aquatic counterpart to dendrochronology. Marine molluscs, which can live for over 500 years, are increasingly providing long-term baseline records of past physical, biological and geochemical variability in marine and aquatic systems over past decades to millennia. Given the annual nature of the growth rings, it is possible to apply dendrochronology derived statistical techniques to construct absolutely dated growth increment width chronologies that facilitate the extension of the sclerochronology records beyond the life span of a single individual over past centuries to millennia.

David’s current research projects involve the development and application of annually resolved absolutely-dated records of past physical, biological and geochemical variability from the North Atlantic, the North Pacific and the Arctic Ocean to investigate local to hemispheric scale connections between marine, atmosphere and cryosphere climate dynamics on inter-annual to millennial timescales. This work involves the application of integrated multi-proxy techniques incorporating sclerochronologies, marine sediment cores, dendrochronologies and ice cores.




Halloran, P.R., Hall, I.R., Menary, M., Reynolds, D.J., Scourse, J.D., Screen, J., Bozzo, A., Dunstone, N., Phipps, S., Schurer, A. P. Sueyoshi, T. and Zhou, T. Natural drivers of multidecadal Arctic sea ice variability over the last millennium. Science Advances.


Foltz, A., Williams, C., Gerson, S.A., Reynolds, D.J, Pogoda S., Begum T. and Walton, S.P., 2019. Game developers’ approaches to communicating climate change. Frontiers. DOI 10.3389/fcomm.2019.00028.

Moffa Sanchez, P., Chamarro-Moreno, E., Reynolds, D.J., Ortega, P., Cunningham, L., Halfar, J., Jonkers, L., Junclaus, J., Perner, K. and Wanamaker, A.D. Jr. 2019. Oceanographic variability in the North Atlantic and Nordic Seas over the Common Era: A review. Paleoceanography and Paleoclimatology. DOI 10.1029/2018PA003508

Reynolds, D.J., Slater, S. and Hall, I.R. 2019. Evaluating the application of carbon isotope records derived from the marine bivalve Glycymeris glycymeris in palaeoenvironmental analyses. Palaeogeography, Palaeoclimatology, Palaeoecology, 523, 48-61, DOI 10.1016/j.palaeo.2019.03.005.

Black, B.A., Andersson, C., Butler, P.G., Carroll, M., DeLong, K.L., Reynolds, D.J., Schöne, B.R., Scourse, J., van der Sleen, P., Wanamaker, A.D. and Witbaard, R. 2019. The revolution of crossdating in marine palaeoecology and palaeoclimatology.  Biology Letters, 15, DOI 10.1098/rsbl.2018.0665


Reynolds D.J., Hall, I.R.,Slater, S.M., Mette, M., Wanamaker, A.D., Scourse, J.D., Garry, F.K., and Halloran, P.R.2018.  Isolating and reconstructing key components of North Atlantic Ocean variability from a sclerochronological spatial network. Paleoceanography and Paleoclimatology, DOI:10.1029/2018PA003366.

Eiriksson, J., Butler, P., Scourse, J.D., Reynolds, D.J, and Símonarson, L.A., 2018. Hafrún – the longest-lived animal known to science. Natturufraedingurinn.


Reynolds D.J., Hall, I.R., Scourse, J.D., Richardson, C.A., Wanamaker, A.D., and Butler, P.G., 2017. Biological and climate controls on North Atlantic marine carbon dynamics over the last millennium: Insights from an absolutely-dated shell based record from the North Icelandic Shelf. Global Biogeochemical Cycles, DOI: 10.1002/2017GB005708

Reynolds, D.J., Hall, I.R.,Slater, S., Scourse, J.D., Halloran, P.R., and Sayer, M.D.J, 2017. Reconstructing past seasonal to multi-centennial scale NE Atlantic Ocean variability using the long-lived marine bivalve mollusc Glycymeris glycymeris. Palaeoceanography 32, 1153-1173, DOI: 10.1002/2017PA003154

Reynolds, D.J., Richardson, C.A., Scourse, J.D., Butler, P.G., Hollyman, P., Roman-Gonzales, A., and Hall, I.R. 2017. Reconstructing North Atlantic marine climate variability using an absolutely-dated sclerochronological network. Palaeogeography, Palaeoclimatology, Palaeoecology 465(B) 333-346, DOI: 10.1016/j.palaeo.2016.08.006.

Román-González, A., Scourse, J., Butler, P., Reynolds, D.J, Richardson, C., Peck, L., Brey, T. and Hall, I. 2017. Analysis of ontogenetic growth trends in two marine Antarctic bivalves Yoldia eightsi and Laternula elliptica: implications for sclerochronology. Palaeogeography, Palaeoclimatology, Palaeoecology 465(B), 300-306, DOI: 10.1016/j.palaeo.2016.05.004.


Reynolds, D.J., Scourse, J.D., Halloran, P.R.Nederbragt, A., Wanamaker, A.D., Butler, P.G., Richardson, C.A., Heinemeier, J., Eiríksson, J., Knudsen, K.L.and Hall, I.R.. 2016. Annually resolved North Atlantic marine climate over the last millennium. Nature Communications 7, DOI: 10.1038/ncomms13502.


Bušelić, I., Peharda, M., Reynolds, D.J., Butler, P.G, Román González,A., Ezgeta-Balić, D., Vilibić, I., Grbec, B., Hollyman, P., Richardson, C.A. 2015. Glycymeris bimaculata (Poli, 1795) – a new sclerochronological archive for the Mediterranean? Journal of Sea Research, 95, 139–148.


Reynolds, D.J., Richardson, C.A., Scourse, J.D., Butler, P.G., Wanamaker, A.D. Jr, Ridgway, I., Sayer, M.D.J. and Gulliver, P. 2013. The potential of the marine bivalve mollusc Glossus humanus (L.) as a sclerochronological archive. The Holocene, 23, 1709-1718.

Reynolds, D.J., Butler, P.G., Williams, S.M., Scourse, J.D., Richardson, C.A. Wanamaker, A.D. Jr., Austin, W.E.N., Cage, A.G. and Sayer, M.D.J. 2013. A multiproxy reconstruction of Hebridean (NW Scotland) spring sea surface temperatures between AD 1805 and 2010. Palaeogeography, Palaeoclimatology, Palaeoecology, 386, 275–285, DOI 10.1016/j.palaeo.2013.05.029.

Brocas W. M., Reynolds D. J., Butler P.G., Richardson C.A., Scourse J.D., Ridgway I. D. And Ramsay K. 2013. The dog cockle, Glycymeris glycymeris (L.), a new annually-resolved sclerochronological archive for the Irish Sea. Palaeogeography, Palaeoclimatology, Palaeoecology, 373, 133-140.

Butler, P. G., Wanamaker Jr, A. D., Scourse, J. D., Richardson, C. A. Reynolds, D. J. 2013. Variability of marine climate on the North Icelandic Shelf in a 1,357-year crossdated Arctica islandica chronology. Palaeogeography, Palaeoclimatology, Palaeoecology, 373, 21-30.


Butler, P.G., Wanamaker, A.D., Jr., Scourse, J.D., Richardson, C.A., Reynolds, D.J. 2011. Long-term stability of δ13C with respect to biological age in the aragonite shell of mature specimens of the bivalve mollusk Arctica islandica. Palaeogeography, Palaeoclimatology, Palaeoecology, 302: 21-30.

Reynolds, D.J. 2011. Stable isotope analysis of biogenic carbonate within the internal growth increments of the marine bivalve Glycymeris glycymeris (L.). Quaternary News, 123.

Ridgway, I.D.  Richardson, C.A.  Scourse, J.D. Butler P.G. And Reynolds, D.J. 2011. The population structure and biology of the ocean quahog, Arctica islandica, in Belfast Lough, Northern Ireland. Journal of the Marine Biological Association of the United Kingdom, Doi:10.1017/S0025315411000154.


Wanamaker Jr, A. D., Baker, A. Butler, P.G. Richardson, C.A. Scourse, J.D. Ridgway, I., Reynolds, D.J. 2009. A novel method for imaging internal growth patterns in marine mollusks: A fluorescence case study on the aragonitic shell of the marine bivalve Arctica islandica (Linnaeus) Limnol. Oceanogr.: Methods 7, 673–681. 


Wanamaker, A.D. Jr., Scourse, J.D., Richardson, C.A. Butler, P.G.  Reynolds D.J. and Ridgeway, I. 2008. Absolute chronologies from the ocean: Records from the longest-lived, non-colonial animals on Earth. PAGES News, 16(3).